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While working with Angus, Dr. Sereno and Jeff Ogradnik noticed that the sole of the forefoot was much larger than the sole of the hind foot. The difference was nearly 1.5 times - the same difference they measured for the weight of the limb supported. That means that each sole is designed to carry the same load per unit area. Footprint size, therefore, is a good estimator for the amount of total body weight carried by a limb.
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